Channa gachua sebelumnya dianggap menyebar luas, bahkan hingga ke Asia Timur dan Indonesia. Berdasarkan banyak kajian genetika dan filogeografi, kelompok C. gachua itu kini telah dibeda-bedakan, paling tidak, atas sekitar 20-an spesies, termasuk yang belum dideskripsi.[4][5][6] Kelompok ikan ini, dengan perkecualian satu spesies, memiliki ciri bersama yaitu pola belang-belang konsentris pada pangkal sirip dadanya.[6]
Di wilayah Srilanka, C. gachua sangat mirip dan dapat tertukar dengan Channa orientalis yang endemik di negeri itu. Sebagian ahli malah menganggapnya bersinonim.[3]
Sementara itu di Burma, Channa harcourtbutleri memiliki bentuk tubuh serupa dan hanya dapat dibedakan melalui pengamatan yang teliti.[7]
C. gachua (sensu stricto) kini diketahui hanya menyebar luas di Asia Selatan. Sedangkan takson serupa yang banyak terdapat di Indonesia bagian barat sekarang dikenal sebagai C. limbata.[4][5][8]
Kelompok Gachua
Beberapa jenis yang termasuk kelompok Gachua, di antaranya:
Channa punctata(Bloch, 1793:139).[29] Asia Selatan, Asia Tenggara, hingga ke Asia Timur: Afghanistan, Pakistan, India, Srilangka, Bangladesh, Burma, Thailand, Malaysia dan Cina; diintroduksi ke negeri-negeri lainnya.
Channa quinquefasciataPraveenraj, Uma, Knight, Moulitharan, Balasubramanian, Bineesh, & Bleher, 2018:143.[30] India utara: aliran S. Brahmaputra: DAS S. Torsa.
Channa raraBritz, Dahanukar, Anoop, & Ali, 2019:590.[5] India, Maharashtra, DAS Jagbudi.
Channa stewartii(Playfair, 1867:14).[32] India (Benggala Barat, Assam), Bangladesh, Bhutan dan Nepal, mungkin juga lebih jauh ke timur.
Channa stiktosLalramliana, Knight, Lalhlimpuia, & Singh, 2018:167.[33] India timurlaut, Mizoram, DAS Kaladan.
Channa torsaensisDey, Nur, Chowdhury, Sarkar, Kosygin, & Barat, 2019:498.[34] India: Benggala Barat, DAS Torsa.
Referensi
^Hamilton, F. (1822). An account of the Fishes found in the River Ganges and its branches. p. 68. Edinburgh : Printed for Archibald Constable & Co. Plate XXI.
^ abRüber, L., H.H. Tan, & R. Britz (2019). "Snakehead (Teleostei: Channidae) diversity and the Eastern Himalaya biodiversity hotspot". J. Zoological Systematics and Evolutionary Research58:356–386. https://doi.org/10.1111/jzs.12324 (laman ResearchGate)
^ abcBritz, R., N. Dahanukar, V.K. Anoop, & A. Ali (2019). "Channa rara, a new species of snakehead fish from the Western Ghats region of Maharashtra, India (Teleostei: Anabantiformes: Channidae)". Zootaxa, 4683:589–600. https://doi.org/10.11646/zootaxa.4683.4.8
^ abSudasinghe, H., R. Pethiyagoda, M. Meegaskumbura, K. Maduwage, & R. Britz (2020). "Channa kelaartii, a valid species of dwarf snakehead from Sri Lanka and southern peninsular India (Teleostei: Channidae)". Vertebrate Zoology70(2):157-170 (April 2020). https://doi.org/10.26049/VZ70-2-2020-05
^Ng, H.H.; Peter K.L. Ng, & R. Britz (1999). "Channa harcourtbutleri (Annandale, 1918): a valid species of snakehead (Perciformes: Channidae) from Myanmar". J. South Asian Nat. Hist., 4: 57-63. (laman ResearchGate)
^Conte-Grand, C., R. Britz, N. Dahanukar, R. Raghavan, R. Pethiyagoda, H.H. Tan, R.K. Hadiaty, N.S. Yaakob, & L. Rüber (2017). "Barcoding snakeheads (Teleostei, Channidae) revisited: discovering greater species diversity and resolving perpetuated taxonomic confusions". PLoS ONE, 12(9): e0184017. https://doi.org/10.1371/journal.pone.0184017
^McClelland, J. (1845). "Description of four species of fishes from the rivers at the foot of the Boutan Mountains". Calcutta Journal of Natural History, 5(18):274-282. (as Ophicephalus amphibeusp. 275)
^ abMusikasinthorn, P. (2000). "Channa aurantimaculata, a new channid fish from Assam (Brahmaputra River basin), India, with designation of a neotype for C. amphibeus (McClelland, 1845)". Ichthyological Research, 47(1):27-37. (laman Academia).
^Britz, R. (2013). "Channa andrao, a new species of dwarf snakehead from West Bengal, India (Teleostei: Channidae)". Zootaxa, 3731(2):287-294. (abstrak).
^Lalhlimpuia, D.V., S. Lalronunga, & Lalramliana (2016). "Channa aurantipectoralis, a new species of snakehead from Mizoram, north-eastern India (Teleostei: Channidae)". Zootaxa, 4147(3):343-350. (abstrak).
^Praveenraj, J., A. Uma, N. Moulitharan, & H. Bleher (2018). "Channa bipuli, a new species of snakehead (Teleostei: Channidae) from Assam, northeast India". aqua, International Journal of Ichthyology, 24(4):153-166. (abstrakDiarsipkan 2022-04-12 di Wayback Machine.).
^Vierke, J. (1991). "Ein farbenfroher neuer Schlangenkopffisch aus Assam: Channa bleheri spec. nov." Das Aquarium, Magazin für zeitgemässe Vivaristik, 25(259): 20-24.
^Chakraborty, P., K. Yardi, & P. Mukherjee (2020). "Channa brahmacharyi sp. nov., a new species of dwarf snakehead from Meghalaya, north-east India (Teleostei: Channidae)". Species, v. 21(67):101-108.
^Praveenraj, J., A. Uma, N. Moulitharan, & R. Kannan (2019). "Channa brunnea, a new species of snakehead (Teleostei: Channidae) from West Bengal, India". Zootaxa, 4624(1):59-70. (abstrak).
^Chaudhuri, B.L. (1919). "Report on a small collection of fish from Putao (Hkamti Long) on the northern frontier of Burma". Records of the Indian Museum (Calcutta), 16(4; art. 18):271-287. (Channa burmanicap.284, Pl. 22. fig. 4, 4a-b).
^Annandale, N. (1918). "Fish and fisheries of the Inlé Lake". Records of the Indian Museum (Calcutta), 14:33-64. (Ophiocephalus harcourt-butlerip. 54, Pl. 2 fig. 7, Pls. 4 figs. 16-17).
^ abNguyen, V.H. (2011). "Two new species belonging to genus Channa (Channidae, Perciformes) discovered in Ninh Binh Province, Vietnam". Tap Chí Sinh Hoc (Journal of Biology), 33(4):8-17. 15 July 2012. (abstract).
^Cuvier, G. & A. Valenciennes (1831). Histoire naturelle des poissons. Tome septième VII:412 (bagian dari uraian Ophicephalus marginatus); Planche 201. F. G. Levrault, Paris. v. 7: i-xxix + 1-531, Pls. 170-208.
^Praveenraj, J., A. Uma, N. Moulitharan, & S.G. Singh (2019). "A new species of dwarf Channa (Teleostei: Channidae) from Meghalaya, northeast India". Copeia, 107(1):61-70. (laman ResearchGate).
^Geetakumari, K. & W. Vishwanath (2011). "Channa melanostigma, a new species of freshwater snakehead from north-east India (Teleostei: Channidae)". Journal of the Bombay Natural History Society107(3) [2010]:231-235.
^Zhang, C.-G., P. Musikasinthorn, & K. Watanabe (2002). "Channa nox, a new channid fish lacking a pelvic fin from Guangxi, China". Ichthyological Research, 49(2):140-146. (laman ResearchGate).
^Bloch, M.E. & J.G. Schneider (1801). M.E. Blochii, Systema Ichthyologiae Iconibus cx Ilustratum. Post obitum auctoris opus inchoatum absolvit, correxit, interpolavit Jo. Gottlob Schneider, Saxo. Berolini. Sumtibus Auctoris Impressum et Bibliopolio Sanderiano Commissum. i-lx + 1-584, Pls. 1-110. (C. orientalisp.496, Pl. 90.
^ abBritz, R. (2008). "Channa ornatipinnis and C. pulchra, two new species of dwarf snakeheads from Myanmar (Teleostei: Channidae)". Ichthyological Exploration of Freshwaters, 18(4) [for 2007]:335-344. (abstrak).
^Musikasinthorn, P. (1998). "Channa panaw, a new channid fish from the Irrawaddy and Sittang River basins, Myanmar". Ichthyological Research, 45(4):355-362. (abstrak).
^Knight, J.D.M. (2016). "Channa pardalis, a new species of snakehead (Teleostei: Channidae) from Meghalaya, northeastern India". Journal of Threatened Taxa8(3):8583-8589.
^Bloch, M.E. (1793). Naturgeschichte der ausländischen Fische. Berlin. v. 7: i-xiv + 1-144, Pls. 325-360. (Ophicephalus punctatus: Pt.2 v.4 th.7:139, Pt.2 Atlas 4: Pl. 358).
^Praveenraj, J., A. Uma, J.D.M. Knight, N. Moulitharan, S. Balasubramanian, K.K. Bineesh, & H. Bleher (2018). "Channa quinquefasciata, a new species of snakehead (Teleostei: Channidae) from Torsa River, North Bengal, India". aqua, International Journal of Ichthyology, 24(4):141-152. (abstrakDiarsipkan 2022-06-23 di Wayback Machine.).
^Endruweit, M. (2017). "Description of a new dwarf snakehead (Perciformes: Channidae) from western Yunnan". Vertebrate Zoology, v. 67(2):173-178.
^Playfair, R.L. (1867). "On the fishes of Cachar". Proceedings of the Zoological Society of London1867(pt 1; art. 3) (for 10 Jan. 1867):14-17. (Ophiocephalus stewartiip.14, Pl. 3).
^Lalramliana, J.D.M. Knight, D.V. Lalhlimpuia, & M. Singh (2018). "Integrative taxonomy reveals a new species of snakehead fish, Channa stiktos (Teleostei: Channidae), from Mizoram, North Eastern India". Vertebrate Zoology68(2):165-175.
^Dey, A., R. Nur, B.R. Chowdhury, D. Sarkar, L. Kosygin, & S. Barat (2019). "A new ornamental species of snakehead fish (Teleostei: Channidae) from River Torsa of West Bengal, India". Int. J. Pure App. Biosci.6(6):497-503 (2018). doi: http://dx.doi.org/10.18782/2320-7051.7131
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