DOSEN PROFIL LENGKAP

constitutes one of the major subdivisions of the phylum Arthropoda. Chelicerates include the sea spiders, horseshoe crabs, and arachnids (including harvestmen...

to palps or palpi) are the secondary pair of forward appendages among chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs...

(the tetrapulmonates); the arachnids as a whole evolved from aquatic chelicerate ancestors. More than 45,000 extant species have been described, organised...

scorpions). It is a terrestrial adaptation of the book gills seen in aquatic chelicerates such as horseshoe crabs, where the gills are ventral appendages under...

Megachelicerax is an extinct genus of stem-chelicerate that lived in Utah during the middle Cambrian. It contains only a single species; Megachelicerax...

early stage of development, so that chelicerates lack the antennae typical of most arthropods. In fact, chelicerates' only appendages ahead of the mouth...

fully sequenced in 2011, and was the first genome sequence from any chelicerate. T. urticae was originally native only to Eurasia, but has acquired a...

Megacheira as closely related to chelicerates, while others recover them as outside the group containing Chelicerate and Mandibulata as stem-group euarthropods...

Offacolus is an extinct genus of euchelicerate, a group of chelicerate arthropods. Its only species, O. kingi, has been found in deposits from the Silurian...

elements of four arthropod-groups: Insects, myriapods, crustaceans, and chelicerates. Insects are used as the model,[why?] with the novel mouthparts of the...

arthropods are a monophyletic group and are divided into three main clades: chelicerates (including arachnids), pancrustaceans (the paraphyletic crustaceans plus...

distinguished from other groups of arthropods, such as insects, myriapods and chelicerates, by the possession of biramous (two-parted) limbs, and by their larval...

Melbournopterus is a genus of prehistoric chelicerate or brachiopod, known from the Upper Silurian of Australia. It is of uncertain taxonomic placement...

middle Cambrian. They are thought to be some of the oldest members of the chelicerate lineage. Members of Habeliida are characterized by a numbers of features...

today have a well developed series of gnathobases along their limbs. Chelicerates have gnathobases that function in conjunction with chelicerae and in...

close to the arachnid crown group. Lustri et al. (2025) propose that chelicerates belonging to the family Offacolidae were suspension feeders. Bicknell...

generally either considering them stem-group euarthropods, or stem-group chelicerates. The homology of the great appendages to the cephalic appendages of other...

2014). "Three-dimensional reconstruction and the phylogeny of extinct chelicerate orders". PeerJ. 2 e641. Bibcode:2014PeerJ...2.e641G. doi:10.7717/peerj...

(/ˌɛkdɪsoʊˈzoʊə/) is a group of protostome animals, including Arthropoda (insects, chelicerates (including arachnids), crustaceans, and myriapods), Nematoda, and several...

three-segmented chelicerae, suggesting that this type is probably ancestral among chelicerates. The jackknife chelicera is subchelate (with fixed finger much reduced...

Despite their name, they are not crabs or even crustaceans; they are chelicerates, more closely related to arachnids like spiders, ticks, and scorpions...

Merostomata is a class of chelicerate arthropods that contains the extinct Eurypterida (sea scorpions) and the extant Xiphosura (horseshoe crabs). The...

close relatives (Artiopoda), Megacheira (which may be paraphyletic) and chelicerates. Under this proposed classification scheme, Arachnomorpha is considered...

The term telson is widely used for the caudal spine of some chelicerates. The chelicerate telson can be clearly seen in a number of fossil species (like...

Mollisoniida is an extinct order of stem group chelicerates that existed from the mid-Cambrian to the Early Ordovician. The clade is united by the presence...

by Limulus, the horseshoe crab, and there are suggestions that other chelicerates developed their simple eyes by reduction from a compound starting point...

Columbia. It is currently considered to be one of the oldest relatives of chelicerates. Unlike most other Burgess forms, Sanctacaris is not present in Charles...

(two branched) limbs. The clade contains all living arthropods (i.e. chelicerates and mandibulates) as well as several fossil groups that share these characteristics...

"Vestigial organs alter fossil placements in an ancient group of terrestrial chelicerates". Current Biology. 34 (6): 1258–1270.e5. Bibcode:2024CBio...34E1258G...

Chasmataspidids, sometime referred to as chasmataspids, are a group of extinct chelicerate arthropods that form the order Chasmataspidida. Chasmataspidids are probably...

Chelicerata. In spiders those usually contain venom glands, but in other chelicerate allies the chelicerae can be chelate without venom glands. Chelae have...

Lozano-Fernandez, L.; et al. (2019). "Increasing species sampling in chelicerate genomic-scale datasets provides support for monophyly of Acari and Arachnida"...

head, it has been widened to include other living arthropods, such as chelicerates, myriapods, and crustaceans, as well as fossil forms, such as the many...

and molluscs; the terrestrial colonization and evolution of insects, chelicerates, myriapods and tetrapods; and the development of modern flora dominated...

other living arthropods, their traditional classification as a member of chelicerates alongside horseshoe crabs and arachnids has regained wide support in...

the genus is significant by bearing several traits of now-surviving chelicerates, such as purportedly pincer-like mouthparts (chelicerae) and fused ring-like...

hyoliths, brachiopods, sponges, chancelloriids, several problematica and a chelicerate are known from the Emu Bay Shale. The Emu Bay Shale of Kangaroo Island...

Chelicerates reported from the Gull River Formation Genus Species Presence Material Notes Images Lunataspis L. borealis Upper Member, Kingston, Ontario...

as the round-tailed horseshoe crab, is a species of horseshoe crab, a chelicerate arthropod found in tropical marine and brackish waters of India, Bangladesh...

subdivided into two tagmata, much like the prosoma and opisthosoma of chelicerate arthropods. Typically, the front part shows no external segmentation...

between trilobitomorphs and chelicerates, while some also interpreted it as a sister group of crustaceans or chelicerates as well. The suggested close...

(2014), "Three-dimensional reconstruction and the phylogeny of extinct chelicerate orders", PeerJ, 2: e641, doi:10.7717/peerj.641, PMC 4232842, PMID 25405073...

as synziphosurine, a paraphyletic group of horseshoe crab-like fossil chelicerate arthropods, while some studies compare its morphology to an artiopod...

polyphyletic (of multiple origins). Current understanding of probable chelicerate relationships, after Sharma and Gavish-Regev (2025): However, a few phylogenomic...

(2014). "Three-dimensional reconstruction and the phylogeny of extinct chelicerate orders". PeerJ. 2 e641. doi:10.7717/peerj.641. PMC 4232842. PMID 25405073...

The crustacean Cherax warsamsonicus The amplypygid chelicerate Phrynus asperatipes In chelicerates and crustaceans, the cephalothorax is derived from...

Weinbergina is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Fossils of the single and type species, W. opitzi, have been...

and hexapods. The evidence for this relationship between myriapods and chelicerates derives from comparisons of nuclear ribosomal RNA genes, mitochondrial...

animals include the cuticle skeletons shared by arthropods (insects, chelicerates, myriapods and crustaceans) and tardigrades, as well as the skeletal...

1. Liestal: Lüdin. Giribet, Gonzalo (March 2018). "Current views on chelicerate phylogeny—A tribute to Peter Weygoldt". Zoologischer Anzeiger. 273: 7–13...

Middle Cambrian, thought to be one of the earliest known relatives of chelicerates. Its fossils have been found in the Burgess Shale in British Columbia...

including cnidaria, echinoderms, molluscs, tunicates, sponges, and some chelicerates. Moving by pseudopodia, amebocytes can manifest as blood cells or play...

thorax (θώραξ, thórax). This fusion of the head and thorax is seen in chelicerates and crustaceans; in other groups, such as the Hexapoda (including insects)...

Foelix (1996), p. 37. Firstman, Bruce (1973). "The Relationship of the Chelicerate Arterial System to the Evolution of the Endosternite". The Journal of...

The pygidium (pl.: pygidia) is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites....

(2014). "Three-dimensional reconstruction and the phylogeny of extinct chelicerate orders". PeerJ. 2 e641. doi:10.7717/peerj.641. PMC 4232842. PMID 25405073...

(2026). "A chelicera-bearing arthropod reveals the Cambrian origin of chelicerates". Nature. 652 (8111): 931–937. Bibcode:2026Natur.652..931L. doi:10...

(2014). "Three-dimensional reconstruction and the phylogeny of extinct chelicerate orders". PeerJ. 2 e641. doi:10.7717/peerj.641. PMC 4232842. PMID 25405073...

worms Pterobranchs Nematodes Chromadorea Enoplea Secernentea Arthropods Chelicerates inc. arachnids Myriapods Pancrustaceans inc. hexapods Rotifera Bdelloidea...

Sanctacaris as stem-group chelicerates. Jago, James B.; García-Bellido, Diego C.; Gehling, James G. (2016). "An early Cambrian chelicerate from the Emu Bay Shale...

The Big Hill Formation is a geologic formation in Michigan. It preserves fossils dating back to the Ordovician period. A fossiliferous site on the Stonington...

Mycteroptidae or Mycteropidae are a family of eurypterids, a group of extinct chelicerate arthropods commonly known as "sea scorpions". The family is one of three...

PMC 11770530. PMID 38623697. Giribet G (March 2018). "Current views on chelicerate phylogeny—A tribute to Peter Weygoldt". Zoologischer Anzeiger. 273: 7–13...

Setapedites is an extinct genus of Ordovician offacolid chelicerates from the Fezouata Formation of Morocco. The genus contains a single species, Setapedites...

Megarachne is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Megarachne have been discovered in deposits of Late Carboniferous...

found in all arachnids (with the exception of some Acari), and in other chelicerates, such as horseshoe crabs. The coxal gland is thought to be homologous...

Rhenopteroidea is an extinct superfamily of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". The superfamily is one...

crab, is a species of horseshoe crab, a kind of marine and brackish chelicerate arthropod. It is found in the Gulf of Mexico and along the Atlantic coast...

Smotrychaspis is a genus of chelicerate arthropod from the Silurian aged Ustya Formation of Ukraine. Representing part of the paraphyletic "synziphosurine"...

non-crustacean arthropods have a single pair of antennae, except for the chelicerates and proturans, which have none. Crustaceans bear two pairs of antennae...

These particular appendages give it an appearance somewhat comparable to chelicerate arthropods such as arachnids and horseshoe crabs. The facehugger is a...

and head appendages, concluded that it was neither a trilobite, nor a chelicerate, nor a crustacean. Marrella is one of several unique arthropod-like organisms...

book gills, and that function in locomotion and gas exchange. In most chelicerates the opisthosomal limbs are greatly reduced and persist only as specialized...

Dekatriata is a clade of planatergan chelicerates including the groups Arachnida, Chasmataspidida, Eurypterida and additionally two stem-genera Winneshiekia...

and leeches (Annelida) Insects, springtails, crustaceans, myriapods, chelicerates (Arthropoda) Chitons, snails, slugs, bivalves, tusk shells, cephalopods...

(breathing) on land provided the gills are kept moist. In some terrestrial chelicerates species (tetrapulmonates such as spiders and scorpions), their ancestral...

with Tetrapulmonata make up the clade Arachnopulmonata. Cladogram of chelicerate relationships, after Sharma and Gavish-Regev (2025): The internal phylogeny...

controversial phylogenetic position either as stem chelicerates or a distinct lineage basal than both chelicerates and mandibulates. Within megachierans, Yohoia...

chains in crustaceans are about 660 amino acid residues long, and in chelicerates they are about 625. In the large complexes there is a variety of variant...

approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids...

Mandibulata and comprise all living arthropods except arachnids and other chelicerates. Unlike chelicerae, mandibles are often used to chew food. Insect mandibles...

Megalograptidae are a family of eurypterids, an extinct group of aquatic chelicerate arthropods commonly known as "sea scorpions". The megalograptids were...

(2017-12-21). "Mandibulate convergence in an armoured Cambrian stem chelicerate". BMC Evolutionary Biology. 17 (1): 261. Bibcode:2017BMCEE..17..261A...

them, but is not up to date for all taxa. Arachnids are eight-legged chelicerate arthropods with pedipalps and bodies divided into two tagmata. The most...

arthropods is uncertain. Some scholars consider them closely related to chelicerates (which include horseshoe crabs, sea spiders, and arachnids) as part of...

Dibasterium is an extinct genus of euchelicerate, a group of chelicerate arthropods. Fossils of the single and type species, D. durgae, have been discovered...

most conspicuous exceptions are the Pycnogonida, which are probably chelicerates. In entomology, the labrum amounts to the "upper lip" of an insect mouth...

posits that artiopods and marrellomorphs are more closely related to Chelicerates. Scholtz, Gerhard; Edgecombe, Gregory (2005-04-27), Koenemann, Stefan;...

(2017-12-21). "Mandibulate convergence in an armoured Cambrian stem chelicerate". BMC Evolutionary Biology. 17 (1): 261. Bibcode:2017BMCEE..17..261A...

description), a group that increasingly evident to be a member of stem-group chelicerates. Within putative habeliids, Dicranocaris might be a close relative of...

arthropods, considered a transitional form between trilobites and true chelicerates. https://web.archive.org/web/20101221002449/http://palaeos...

Montenegro is one of two and the main hotspot for troglobitic fauna in Western Balkans (though fragments of this hotspot reach into the easternmost tips...

was explored by crustaceans but unique within the dominantly carnivore chelicerates. This hypothesis is supported by the morphology of their prosomal exopods...

called the Nerscylla, described in game as a "Temnoceran," based on the Chelicerate subphylum of arthropods, along with its subspecies, the Shrouded Nerscylla...

demersal fishes) and crawl around (e.g. decapod crustaceans, marine chelicerates, octopus, most non-bivalvian molluscs, echinoderms etc.). Many benthic...

member of Sanctacarididae, a family which later grouped under the stem chelicerate order Habeliida. Utahcaris is distinguishable from other sanctacaridids...

Hou, X.; Ma, X.; Edgecombe, G.D.; Strausfeld, N.J. (October 2013). "Chelicerate neural ground pattern in a Cambrian great appendage arthropod". Nature...

unresolved, with some phylogenies recovering Artiopoda as more closely to chelicerates, forming the clade Arachnomorpha, while others recover Artiopoda as more...

Eurypterids are a group of chelicerates that became extinct in the Late Permian. They underwent ecdysis similarly to extant chelicerates, and most fossils are...

by Limulus, the horseshoe crab, and there are suggestions that other chelicerates developed their simple eyes by reduction from a compound starting point...

genus is a basal chelicerate, closer to crown group Chelicerata than members of Habeliida. Thelxiope Arthropoda stem-groupChelicerate Walcott Quarry A...

spanning the Middle Cambrian to the Upper Ordovician. Initially considered chelicerates, modern anatomical comparisons demonstrate that the aglaspidids cannot...

"Vestigial organs alter fossil placements in an ancient group of terrestrial chelicerates". Current Biology. 34 (6): 1258–1270.e5. doi:10.1016/j.cub.2024.02.011...

Kokomopteroidea is an extinct superfamily of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". It is one of four superfamilies...

Winneshiekia is an extinct genus of dekatriatan, a clade of chelicerate arthropods. Fossils of the single and type species, W. youngae, have been discovered...

Barusopterus is a genus of strobilopterid eurypterid that lived in Indiana, US during the Silurian period. This genus contains a single species, Barusopterus...

Pseudoniscidae is an extinct family of synziphosurine chelicerates that lived in the Silurian. Pseudoniscidae is classified inside the clade Planaterga...

'great-appendage' arthropod from the Lower Cambrian of China and homology of chelicerate chelicerae and raptorial antero-ventral appendages". Lethaia. 37 (1):...

stridulating organ present but reduced. They also own thin and elongated chelicerate strikers, which are pallid in color. Their size is also smaller than...

Carcinosomatoidea is an extinct superfamily of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". It is one of the superfamilies...

Synziphosurina is a paraphyletic group of chelicerate arthropods previously thought to be basal horseshoe crabs (Xiphosura). It was later identified as...

26879/1547. Legg, David A. (December 2014). "Sanctacaris uncata: the oldest chelicerate (Arthropoda)". Naturwissenschaften. 101 (12): 1065–1073. Bibcode:2014NW...

Schneiderman HA (December 1970). "Control of molting in mandibulate and chelicerate arthropods by ecdysones". The Biological Bulletin. 139 (3): 520–538....

Bunaia is a genus of Silurian chelicerate arthropod. The genus contains at least one species: Bunaia woodwardi from the Silurian period in Svalbard, Norway...

Eukoenenia spelaea (Palpigradi) Character states and evolution of the chelicerate claws Barnes, Robert D. (1982). Invertebrate Zoology. Philadelphia, PA:...

Megalograptus might have been cannibalistic at times, like many modern chelicerates. Megalograptus was a genus of large megalograptids. The largest species...

Mycteropoidea is an extinct superfamily of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". It is one of four superfamilies...

Eurypterina is one of two suborders of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". Eurypterine eurypterids...

(2014). "Three-dimensional reconstruction and the phylogeny of extinct chelicerate orders". PeerJ. 2: e641. doi:10.7717/peerj.641. ISSN 2167-8359. PMC 4232842...

'great-appendage' arthropod from the Lower Cambrian of China and homology of chelicerate chelicerae and raptorial antero-ventral appendages". Lethaia. 37 (1):...

Legrandella is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Legrandella was regarded as part of the clade Prosomapoda...

Bunodidae is an extinct family of synziphosurine chelicerates that lived in the Silurian. Bunodidae is classified inside the clade Planaterga alongside...

surface by the sternites. In scorpions, the metasoma is the tail. In other chelicerates, such as spiders, the mesosoma is fused with the metasoma to form the...

Xiaoya; Edgecombe, Gregory D.; Strausfeld, Nicholas J. (2013-10-17). "Chelicerate neural ground pattern in a Cambrian great appendage arthropod". Nature...

Limulus is a genus of horseshoe crab, with one extant species, the Atlantic horseshoe crab (Limulus polyphemus). One fossil species is currently assigned...

Limuloides is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Limuloides was regarded as part of the clade Planaterga....

Pagea is a genus of prehistoric eurypterid classified as part of the family Stylonuridae. It contains three species, all from the Devonian (Lochkovian...

"Vestigial organs alter fossil placements in an ancient group of terrestrial chelicerates". Current Biology. "Ancient four-eyed wonder resolves daddy longleg mystery"...

"Phylogeny and systematic position of Opiliones: a combined analysis of chelicerate relationships using morphological and molecular data" (PDF). Cladistics...

Borchgrevinkium is an extinct genus of chelicerate arthropod. A fossil of the single and type species, B. taimyrensis, has been discovered in deposits...

Leanchoilia superlata - megacheiran Megachelicerax - chelicerate Mollisonia symmetrica - mollisoniid chelicerate Perspicaris dilatus hymenocarine Pseudoarctolepis...

close relatives Mollisonia and Corcorania. They are suggested to be stem-chelicerates. "Thelxiope palaeothalassia". Burgess Shale Fossil Gallery. Virtual Museum...

Titanoprosoma is an extinct genus of chelicerate with uncertain affinities. The only known species is T. edgecombei, found in the Serpukhovian Bear Gulch...

Rhenopterus is an extinct prehistoric eurypterid. Fossils of Rhenopterus have been recovered from deposits of Lower Devonian age in Germany. Rhenopterids...

led to a possible affinity -or being a close relative of trilobites or chelicerates or possibly a close relative of both. Because the presumed intestinal...

none of the known marine arthropod groups (trilobites, crustaceans, chelicerates). Yohoia was a tiny animal (7 mm to 23 mm long) with: a head shield;...

megacheiran genus Yawunik, the isoxyid genus Surusicaris, and the basal chelicerate Mollisonia plenovenatrix. Worms have also been found at this site, like...

position; their brain anatomy is more closely related to that of the chelicerates than to any other arthropod. The modern velvet worms form a monophyletic...

Lyall I. (January 2002). "A New Three-Dimensionally Preserved Xiphosuran Chelicerate from the Montceau-Les-Mines Lagerstätte (Carboniferous, France)". Palaeontology...

tome VI, Onychophores - Tardigrades - Arthropodes - Trilobitomorphes - Chélicérates. - Masson et Cie, Paris. Shultz, J.W. (1990). Evolutionary Morphology...

Venustulus is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Venustulus was regarded as part of the clade Prosomapoda...

Eurypterus (/jʊəˈrɪptərəs/ yoo-RIP-tər-əs) is an extinct genus of eurypterid, a group of organisms commonly called "sea scorpions". The genus lived during...

Ciurcalimulus is a genus of xiphosuran from the Silurian-aged Wabash Formation of Indiana, United States. It is one of the earliest known relatives of...

(2014). "Three-dimensional reconstruction and the phylogeny of extinct chelicerate orders". PeerJ. 2 e641. doi:10.7717/peerj.641. ISSN 2167-8359. PMC 4232842...

of Mollisonia appendages in 2019, the group is suggested to be stem-chelicerates. However, a redescription published in July 2024 discovered the appendages...

parisiana A thylacocephalan Limulidae? Gen. et sp. indet. A xiphosuran chelicerate Litogaster L. turnbullensis? A glypheid pleocyemate crustacean Litogaster...

Cheliceratichnus, meaning trace of a chelicerate arthropod, is an ichnogenus erected for a body imprint discovered in the Lower Jurassic East Berlin Formation...

Chasmataspis is a genus of chasmataspidid, a group of extinct aquatic chelicerate arthropods. It was found in the Early Ordovician deposits of Tennessee...

(2014). "Three-dimensional reconstruction and the phylogeny of extinct chelicerate orders". PeerJ. 2 e641. doi:10.7717/peerj.641. PMC 4232842. PMID 25405073...

evolved walking several times, specifically in hexapods, myriapods, chelicerates, tardigrades, onychophorans, and crustaceans. Little skates, members...

lower Ordovician unit right at the top of the formation, containing the chelicerate Esmeraldacaris, as well as some graptolite fauna. This section details...

Bunodes is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Bunodes was regarded as part of the clade Planaterga. Fossils...

known features shared by all genera within the group. Like all other chelicerates, and other arthropods in general, waeringopterid eurypterids possessed...

Retrieved 4 May 2026. Giribet, Gonzalo (1 March 2018). "Current views on chelicerate phylogeny—A tribute to Peter Weygoldt". Zoologischer Anzeiger. 273: 7–13...

rapid growth. Unlike many arthropods, such as insects and crustaceans, chelicerates (the group to which eurypterids like Jaekelopterus belongs, alongside...

"Vestigial organs alter fossil placements in an ancient group of terrestrial chelicerates". Current Biology. 34 (6): 1258–1270.e5. doi:10.1016/j.cub.2024.02.011...

and at least three opsins were present in the ancestor of arthropods; chelicerates and pancrustaceans today possess color vision. Researchers studying the...

close to the crown-group of Euarthropoda, which includes myriapods, chelicerates, insects and crustaceans. Briggs, D.E.G. (1977). "Archived copy" (PDF)...

Camanchia is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Camanchia was regarded as part of the clade Prosomapoda....

Heethoff (28 March 2017), "Storage and release of hydrogen cyanide in a chelicerate ( Oribatula tibialis )", Proceedings of the National Academy of Sciences...

possibly could". As an example, the sea scorpions, an extinct group of chelicerate arthropods encountered by Marven in the Ordovician, were developed and...

Bembicosoma is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Bembicosoma was regarded as part of the clade Planaterga...

Pterygotus is an extinct genus of giant predatory eurypterid, a group of extinct aquatic arthropods. Fossils of Pterygotus have been discovered in deposits...

Eocarcinosoma is a genus of eurypterid, an extinct group of aquatic arthropods. The type and only species of Eocarcinosoma, E. batrachophthalmus, is known...

(2014), "Three-dimensional reconstruction and the phylogeny of extinct chelicerate orders", PeerJ, 2 e641, doi:10.7717/peerj.641, PMC 4232842, PMID 25405073...

Angel, Matarael (マトリエル, Matorieru), resembles an arthropod arachnid chelicerate. Unlike arthropods, it has a single central body, from the top of which...

evolutionary biology Evolution of molluscs Evolution of spiders – Origin from a chelicerate ancestor and diversification of spiders through geologic time Evolution...

"Isolation and sequence analysis of the gene for arginine kinase from the chelicerate arthropod, Limulus polyphemus: insights into catalytically important...

Hunanopterus is a genus of eurypterid, an extinct group of aquatic arthropods. The genus contains one species, the type species H. wangi from the Silurian...

'great-appendage' arthropod from the Lower Cambrian of China and homology of chelicerate chelicerae and raptorial antero-ventral appendages". Lethaia. 37 (1):...

Lunataspis is the oldest known xiphosuran, with three known species all dating from the late Ordovician (latest Sandbian for L. borealis, Katian for L...

Recent studies around 2020s suggest it is possibly a habeliid (stem chelicerate) alongside Dicranocaris and Messorocaris. Briggs, Derek E. G.; Lieberman...

Eusarcana (meaning "true flesh") is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Eusarcana have been discovered in deposits...

three segments called Tagmata and in the case of chelicerates, they only possess two. In chelicerates, these tagmata are called the Prosoma and the Opisthosoma...

Prashant P (2023-02-17). "Dual Functions oflabialResolve the Hox Logic of Chelicerate Head Segments". Molecular Biology and Evolution. 40 (3) msad037. doi:10...

Messorocaris and both are tentatively grouped under Habeliida (stem-group chelicerates). Briggs, Derek E. G.; Lieberman, Bruce S.; Hendricks, Jonathan R.; Halgedahl...

Cyamocephalus is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Cyamocephalus was regarded as part of the clade Planaterga...

Pentecopterus is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils have been registered from the Darriwilian age of the Middle Ordovician...

nymphal stages and an adult stage. Oribatid mites differ from other chelicerate animals such as spiders by the fact that they feed on food particles...

consider it a sister group to Arachnomorpha, which includes trilobites and chelicerates (see cladogram below). Both are grouped under Schizoramia, a clade of...

Anderson, Lyall I. (2002). "A New Three-Dimensionally Preserved Xiphosuran Chelicerate from the Montceau-Les-Mines Lagerstätte (Carboniferous, France)". Palaeontology...

Magnicornaspis is an extinct genus of chelicerate arthropod belonging to the family Corcoraniidae. It was discovered in the Rivière-du-Loup Formation...

Interestingly, Notchia share similarities with some chelicerates, such as a reduced first tergite, although no chelicerate so far has such a short cephalon or similar...

the abdomen, and lunging. Palps are a second pair of appendages on the chelicerates on C. umbratica. Males utilize their ability to reflect UV light as a...

Prolimulus is an extinct genus of arthropods belonging to the order Xiphosura and family Belinuridae. Fossils of this genus were discovered in the Czech...

2005). "A glyptocystitid cystoid affinity for the putative stem group chelicerate (Arthropoda: Aglaspidida or Xiphosura) Lemoneites from the Ordovician...

comparative genomics of ancient invertebrate groups, with emphasis on chelicerate arthropods. He is the director of the Zoological Museum of the University...

controversial phylogenetic position. It was previously thought to be chelicerate-related, but later studies repeatedly suggests it as a member of Artiopoda...

species. Briggs, Derek E. G.; Collins, Desmond (1988). "A Middle Cambrian chelicerate from Mount Stephen, British Columbia" (PDF). Palaeontology. 31 (3): 779–798...

Herefordopterus is a genus of eurypterid, an extinct group of aquatic arthropods. Herefordopterus is classified as the only genus in the family Herefordopteridae...

early cladistic analysis suggested, rather, that it was a stem-group chelicerate. The presence of a distinct anterior sclerite bearing the eyes has been...

Ward C.; Hayashi, Cheryl Y. (June 1998). "The Phylogeny of the Extant Chelicerate Orders". Cladistics. 14 (2): 173–192. doi:10.1006/clad.1998.0061. Ballesteros...

Peltidium is a prodorsal shield found in animals of the subphylum Chelicerata, in the phylum Arthropoda. In some groups (Schizomida, Palpigradi, Solpugida...

1956, available here. ISBN 0-8137-3016-3. --- Part P refers to extinct chelicerates including eurypterids (sea scorpions), xiphosurans (horseshoe crabs)...

chose M. inclemens, and this has become the standard name. Spiders are chelicerates, which differ from other arthropods in that the usual body segments are...

Kiaeria is a genus of chasmataspidid, a group of extinct aquatic arthropods. It was originally classified as a xiphosuran of the monotypic family Kiaeriidae...

the club discovered the first and only known fossil of a prehistorical chelicerate or brachiopod specimen in Heathcote. The specimen was initially identified...

possess large, functional eyes, presumably because these spider-like chelicerates feed on cave-dwelling, light-emitting glowworm larvae Arachnocampa which...

specimens and diversity. A wide variety including crustaceans, trilobites, chelicerates, and less familiar groups like the thylacocephalans and cheloniellids...

Kokomopterus is a genus of prehistoric eurypterid. The genus contains a single species, Kokomopterus longicaudatus, known from the Silurian of Kokomo,...

Alkenopterus is a genus of prehistoric eurypterid. The genus contains two species, A. brevitelson and A. burglahrensis, both from the Devonian of Germany...

Hallipterus is a genus of prehistoric eurypterid classified as part of the family Hardieopteridae. Hallipterus was a gigantic Hardieopterid eurypterid...

Woodwardopterus is a genus of prehistoric eurypterids, colloquially known as sea scorpions, classified as part of the family Mycteroptidae. The genus contains...

Parastylonurus is a genus of prehistoric eurypterid. It is the only genus classified in the family Parastylonuridae and contains two species, P. hendersoni...

1007/BF02988492. Poschmann, Markus; Anderson, Lyall I.; Dunlop, Jason A. (2005). "Chelicerate Arthropods, including the Oldest Phalangiotarbid Arachnid, from the Early...

Animalia (Animals) Phylum Arthropoda (Arthropods) Subphylum Chelicerata (Chelicerates) Class Arachnida (Arachnids) Order Araneae (Spiders) Infraorder Araneomorphae...

Archopterus is a genus of eurypterid in the family Pittsfordipteridae, represented by one species. It was discovered in the Wenchang Formation of Tianfucun...

cartilaginous fish and some homology in insects, lobe finned fish, chelicerates and corals (Figure 22). The following significant taxonomic groups do...

Stylonuroidea is an extinct superfamily of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". It is one of four superfamilies...

Republic, which bears some resemblance to the eurypterids, aglaspidids and chelicerates. It is diagnostic of a small and low-diversity fauna endemic to the area...

Dahms, Hans-Uwe (2011). "Early Post-Embryonic Development of Marine Chelicerates and Crustaceans with a Nauplius" (PDF). Crustaceana. 84 (7): 869–893...

Lyall I. (January 2002). "A New Three-Dimensionally Preserved Xiphosuran Chelicerate from the Montceau-Les-Mines Lagerstätte (Carboniferous, France)". Palaeontology...

from Alken, Germany. It was initially described by Leif Størmer as a chelicerate arthropod possibly belonging to the order Eurypterida. Nowadays, it is...

Slimonia is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Slimonia have been discovered in deposits of Silurian age in South...

The Hardieopteridae are a family of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". The family is one of two...

but based on the medial position of anus (which, in telson-bearing chelicerates, always located at the ventral boundary of abdomen and telson), the latter...

Casterolimulus is an extinct genus of xiphosuran. It is known from the Late Cretaceous Fox Hills Formation of North America, and is believed to have lived...

Xaniopyramis is an extinct genus of xiphosuran, related to the modern horseshoe crab. It lived in the early Carboniferous. Xaniopyramis linseyi Siveter...

Euproops is very similar to that of living horseshoe crabs and other chelicerates. Unlike living horseshoe crabs, many belinurids are thought to have primarily...

Houia is an extinct genus of dekatriatan, a clade of chelicerate arthropods. Fossils of Houia have been discovered in deposits of the Early Devonian period...

Hoplitaspis is a genus of chasmataspidid, an extinct group of aquatic arthropods. Fossils of Hoplitaspis have been discovered in Late Ordovician deposits...

(2017). "The gnathobasic spine microstructure of recent and Silurian chelicerates and the Cambrian artiopodan Sidneyia: Functional and evolutionary implications"...

Liomesaspis is an extinct genus of xiphosuran, related to the modern horseshoe crab. It lived from the late Carboniferous to the Early Permian. Evolution:...

valuable resource for functional and evolutionary genomic studies in chelicerates". GigaScience. 8 (8) giz099. doi:10.1093/gigascience/giz099. PMC 6701490...

arthropods such as trilobites, radiodonts, marrellids, artiopods, and stem-chelicerates. It is part of the larger Balang Formation, and is roughly contemporaneous...

Member/Ellicott Creek Breccia Bunaia B. woodwardi Williamsville A Offacolid Chelicerate. Carcinosoma C. scorpionis Williamsville Renamed as Cruinnopterus Ceratiocaris...

Lanarkopterus is a genus of prehistoric eurypterid with one recognised species, Lanarkopterus dolichoschelus. Lanarkopterus was long seen as a species...

Palaeoscorpius A primitive scorpion Schinderhannes 1 Specimen A radiodont Weinbergina A chelicerate Wingertshellicus An arthropod of uncertain affinities...

pheromones to persist longer. Receiving spiders do so by using chemosensitive chelicerates and may be able to detect specific information about the emitting female...

"Phylogeny and Systematic Position of Opiliones: A Combined Analysis of Chelicerate Relationships Using Morphological and Molecular Data". Cladistics. 18...

which might have reached lengths of 2.6 metres (8.5 ft). Like all other chelicerates, and other arthropods in general, pterygotioid eurypterids possessed...

Vaderlimulus is a genus of horseshoe crab. The species V. tricki lived in the Olenekian age Triassic period approximately 245 million years ago North America...

Bassipterus ("wing from Bass") is a genus of eurypterid, an extinct group of aquatic arthropods. Bassipterus is classified as part of the family Adelophthalmidae...

Clarkeipterus is a genus of prehistoric eurypterid classified as part of the family Dolichopteridae. The genus contains two species, C. otisius and C....

protruding through their cuticle ("skin"). Unlike insects, spiders and other chelicerates do not have antennae. Most species of jumping spiders appear to be cursorial...

Mesolimulus is an extinct genus of horseshoe crab. The best known examples are found in Solnhofen limestone near Solnhofen, Bavaria, Germany. Originally...

Cruinnopterus is a genus of eurypterid, an extinct group of aquatic arthropods. The genus contains two species, the type species C. scorpioides from the...

Vinetopterus is a genus of prehistoric eurypterid from the Devonian period in Europe classified as part of the Moselopteridae family. The genus contains...

Campylocephalus is a genus of eurypterid, a group of extinct aquatic arthropods. Fossils of Campylocephalus have been discovered in deposits ranging from...

A.; Selden, Paul A. (1998). "The early history and phylogeny of the chelicerates". Arthropod Relationships. The Systematics Association Special Volume...

Nanahughmilleria ("dwarf Hughmilleria") is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Nanahughmilleria have been discovered...

Hibbertopterus is a genus of eurypterid, a group of extinct marine arthropods. Fossils of Hibbertopterus have been discovered in deposits ranging from...

Lamontopterus is a genus of prehistoric eurypterid classified within the family Kokomopteridae. It contains one species, Lamontopterus knoxae, from the...

Hastimima is a genus of Brazilian eurypterid, a group of extinct marine arthropods. Fossils of Hastimima have been discovered in Permian coal deposits...

Austrolimulus fletcheri is an extinct xiphosuran, related to the modern horseshoe crab. The species is only known from the Middle Triassic-aged strata...

Belinurus is an extinct genus of arthropods belonging to the order Xiphosura. It is part of the family Belinuridae, in the infraorder Belinurina. There...

Guangyuanolimulus is an extinct genus of limulid xiphosuran that lived during the Permian-Triassic extinction event. Guangyuanolimulus shangsiensis fossils...

Stoermeropterus is a genus of prehistoric eurypterid from the Silurian period in Europe and North America classified as part of the Moselopteridae family...

Cambropycnogon is an extinct genus of pycnogonid arthropods. The genus contains a single species, Cambropycnogon klausmuelleri, represented by presumed...

VI (1949) : Onychophores, Tardigrades, Arthropodes, Trilobitomorphes, Chélicérates. Tome VII, Fascicule 2 (1996) : Crustacés. Tome VIII, Fascicule 1 (1973) :...

Diego C. García-Bellido; James G. Gehling (2016). "An early Cambrian chelicerate from the Emu Bay Shale, South Australia". Palaeontology. 59 (4): 549–562...

Belinuropsis is a dubious genus of prehistoric arthropod which contains one species, B. wigudensis, from the Silurian period of New Brunswick, Canada....

Soligorskopterus is a genus of eurypterid, a group of extinct aquatic arthropods. Fossils of Soligorskopterus have been discovered in deposits from the...

Drepanopterus is an extinct genus of eurypterid and the only member of the family Drepanopteridae within the Mycteropoidea superfamily. There are currently...

Mycterops is a genus of prehistoric eurypterid of the family Mycteroptidae. Mycterops lived during the Carboniferous period in Europe and North America...

The Kokomopteridae are a family of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". The family is one of two...

Ciurcopterus is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Ciurcopterus have been discovered in deposits of Late Silurian...

the Early to Middle Silurian of Lesmahagow, Scotland. Like all other chelicerates, and other arthropods in general, slimonid eurypterids possessed segmented...

Pasternakevia is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Pasternakevia was regarded as part of the clade Planaterga...

Kasibelinurus (meaning "sibling of dart tail" for its perceived relation to Belinurus at the time) is a genus of xiphosuran from the Devonian. It is part...

Stylonuridae is a family of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". The family is one of two families...

Haifengella corona is the only known species of the extinct arthropod genus Haifengella, described in 2014 from the Early Cambrian Chengjiang Lagerstätte...

Salteropterus is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Salteropterus have been discovered in deposits of Late Silurian...

Attenborolimulus is an extinct genus of austrolimulid horseshoe crab with one known species: Attenborolimulus superspinosus. This genus is known from the...

Terropterus is a genus of eurypterid, an extinct group of aquatic arthropods. The type and only species of Terropterus, T. xiushanensis, is known from...

Strobilopterus is a genus of prehistoric eurypterid of the family Strobilopteridae. The genus contains four species, two from the Devonian of Wyoming,...

Stylonurella is a genus of prehistoric eurypterid. It is the only genus classified in the Stylonurellidae family and contains a single valid species, S...

Pentapantopus is a genus of fossil pycnogonid (sea spider). The only known species is Pentapantopus vogteli from the Hunsrück Slate of Germany. This sea...

Flagellopantopus is an extinct genus of pycnogonid (sea spider) arthropod known from the lower Devonian aged Hunsrück Slate. A single species is currently...

Acutiramus is a genus of giant predatory eurypterid, an extinct group of aquatic arthropods. Fossils of Acutiramus have been discovered in deposits of...

at 2.5 metres and Acutiramus bohemicus at 2.1 metres. Like all other chelicerates, and other arthropods in general, pterygotid eurypterids possessed segmented...

Mixopterus is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Mixopterus have been discovered in deposits from Late Silurian...

brachiopods (such as Orbiculoidea), orthoconic nautiloids, naraoiid chelicerates, conodonts and other organisms, O. augusti would have been a top predator...

Dorfopterus is a genus of extinct invertebrate, possibly an eurypterid. Only one fossil of the single and type species, D. angusticollis, has been discovered...

single plate located in the opisthosoma of non-arachnid dekatriatan chelicerates such as eurypterids, chasmataspidids and the genus Houia. The metastoma...

Anderella is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Anderella was regarded as part of the clade Prosomapoda....

Victalimulus mcqueeni is an extinct horseshoe crab, from Early Cretaceous (Aptian) Koonwarra Fossil Beds in eastern Victoria, Australia. The holotype and...

Buffalopterus is a prehistoric eurypterid from the Silurian-aged Bertie Formation of New York and Ontario. The genus contains one species, B. pustulosus...

Corcorania is a genus of mollisoniid arthropod from the early Lancefieldian of Australia. It contains one species, Corcorania trispinosa. Corcorania has...

protruding through their cuticle ("skin"). Unlike insects, spiders and other chelicerates do not have antennae. A Portia can sense vibrations from surfaces, and...

Moselopterus is a genus of prehistoric eurypterid from the Devonian period in Europe. The genus contains two species, M. ancylotelson from Germany and...

Euproops is an extinct genus of xiphosuran, related to the modern horseshoe crab. It lived during the Carboniferous Period. The Treatise on Invertebrate...

Eysyslopterus is a genus of eurypterid, an extinct group of aquatic arthropods. Eysyslopterus is classified as part of the family Waeringopteridae. One...

Brachyopterella is a genus of prehistoric eurypterid classified as part of the Brachyopterellidae family. Two species, both from the Silurian period, are...

smell, taste, touch and vibration. Unlike insects, spiders and other chelicerates do not have antennae. While most jumping spiders walk quickly, in a stop-go...

Carcinosoma (meaning "crab body") is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Carcinosoma are likely restricted to deposits...

Ctenopterus is a genus of prehistoric eurypterid of the family Stylonuridae. It contains only one species, Ctenopterus cestrotus from the Early Silurian...

small to support chewing mandibles. It also does not match closely with chelicerates, as their head muscles are also too specialised for Keurbos to belong...

Hughmilleria is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Hughmilleria have been discovered in deposits of the Silurian...

Haliestes is a genus of sea spider (pycnogonid) from the Silurian aged Coalbrookdale Formation of England. It contains a single species, Haliestes dasos...

Mymarommatidae. Protorthoptera and Palaeodictyoptera also occur. Frequent chelicerates (arthropods) include trace and body fossils of xiphosurid merostomes...

Grossopterus is a genus of prehistoric eurypterid classified as part of the family Waeringopteridae. The genus contains two species, G. inexpectans from...

Wiedopterus is a genus of eurypterid, an extinct group of aquatic arthropods. The type and only species of Wiedopterus, W. noctua, is known from deposits...

Stylonuroides is a genus of prehistoric eurypterid. The genus contains a single species, S. dolichopteroides from the Silurian of Ringerike, Norway. Stylonuroides...

of the tail plate. A possible synapomorphy of the appendages with the chelicerates is purely speculative. A later cladistic analysis resolved Kiisortoqia...

Brachyopterus was one of the earliest known eurypterid genera, having been recovered from Middle Ordovician deposits in Montgomeryshire, Wales. Though...

Ortega-Hernández, Javier (2020-04-09). "Revision of the mollisoniid chelicerate(?) Thelxiope, with a new species from the middle Cambrian Wheeler Formation...

Hardieopterus is a genus of prehistoric eurypterid classified within the family Hardieopteridae. The genus was previously considered to contain four species...

Dvulikiaspis is a genus of chasmataspidid, a group of extinct aquatic arthropods. Fossils of the single and type species, D. menneri, have been discovered...

Laurieipterus is a genus of a eurypterid classified as part of the family Stylonuridae. It contains one species, L. elegans from the Early Silurian of...

published by Jason A. Dunlop and Paul A. Selden (1998) is based on the major chelicerate groups (in bold, Aglaspida, Eurypterida and Xiphosurida, Scorpiones and...

parva The youngest known "synziphosurine" ( a group of xiphosuran-like chelicerates) Bairdops B. beargulchensis A possibly paraphyletic "palaeostomatopod"...

bromalites, with a number of arthropods (primarily various crustaceans and chelicerates), agnathans, algae, gastropods and brachiopods making up the rest of...

Waeringopterus is a genus of prehistoric eurypterids from the Silurian of North America. The genus contains two species, W. apfeli from the Syracuse and...

Unionopterus is a genus of eurypterid, an extinct group of aquatic arthropods commonly known as "sea scorpions". Fossils have been registered from the...

the past have been thought to represent an entirely distinct order of chelicerates. Fossils of the family first appear in deposits of the Middle Devonian...

inhabit the shore areas for calmer waters. This species is a marine chelicerate arthropod, rather than a crab, and has a strange similarity to spiders...

Stylonurus is a genus of prehistoric eurypterid of the family Stylonuridae. The genus contains three species: Stylonurus powriensis from the Devonian of...

Erieopterus is a genus of prehistoric eurypterid found in Silurian to Devonian-aged marine and freshwater strata of North America. The genus contains three...

Adelophthalmus is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Adelophthalmus have been discovered in deposits ranging in...

Palaeopantopus is a monotypic genus of fossil pycnogonid (sea spider), known only by one species, Palaeopantopus maucheri from the Devonian Hunsrück Slate...

discussion, with some studies classifying them as either stem-group chelicerates, or stem-group deuteropods. Parry et al., 2024 conducted multiple phylogenetic...

protruding through their cuticle ("skin"). Unlike insects, spiders and other chelicerates do not have antennae. A Portia can sense vibrations from surfaces, and...

structures with those of other crustaceans, and parallels drawn instead with chelicerates. However, other workers have interpreted the gills as similar to those...

Ruedemannipterus is a genus of prehistoric eurypterid classified as part of the family Dolichopteridae. The genus contains one species, R. stylonuroides...

Forfarella is a genus of chasmataspidid, a group of extinct aquatic arthropods. Fossils have been discovered in deposits of the Early Devonian period....

yielded numerous early land plants, arthropods and vertebrates. Among the chelicerates, to which the eurypterids belong, the Early Devonian deposits have yielded...

enlarged. Another example would be Necrogammarus, an extinct genus of chelicerate eurypterid only known by its subcapitulum and attached pedipalp. In this...

probably of digestive enzymes similar to those in modern crustaceans or chelicerates. Gishlick & Fortey (2023) describe a specimen of Walliserops trifurcatus...

Paraeurypterus is a genus of prehistoric eurypterid from the Late Ordovician period. The genus contains one species, P. anatoliensis, known from Şort Tepe...

Vicissicaudata is more closely related to other arthropod groups like chelicerates than to trilobitomorphs. Cladogram after McCoy et al. 2025: Cladogram...

Kiaeropterus is a genus of prehistoric eurypterid classified as part of the Brachyopterellidae family. Two species, both from the Silurian period, are...

Dolichopterus is a genus of prehistoric sea scorpions, arthropods in the order Eurypterida. Fossils of Dolichopterus have been discovered in deposits ranging...

Rhinocarcinosoma is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Rhinocarcinosoma have been discovered in deposits ranging...

Polystomurum is a Devonian genus from Siberia, Russia (then the Soviet Union). It was initially described by Nestor Novojilov in 1958 as a eurypterid with...

Echinognathus is a genus of eurypterid, an extinct group of aquatic arthropods. The type and only species of Echinognathus, E. clevelandi, is known from...

likely made by eurypterids or larger conodonts. Other fossils include chelicerates, bivalved crustaceans, algae, linguloid brachiopods, a single gastropod...

cuticle is punctured using a pair of jaws homologous to the chelicerae of chelicerates. These are not mandibles, because they belong to the second body segment...

Erettopterus is a genus of large predatory eurypterid, an extinct group of aquatic arthropods. Fossils of Erettopterus have been discovered in deposits...

Crustaceans, chelicerates, molluscs Other invertebrates American lobster orange-footed sea cucumber rock crab Forbes' sea star jonah crab Atlantic purple...

from the known marine arthropod groups (trilobites, crustaceans and chelicerates) in the structure of its legs and gills, and above all in the number...

published by Jason A. Dunlop and Paul A. Selden (1998) is based on the major chelicerate groups (in bold, Aglaspida, Eurypterida and Xiphosurida, Scorpiones and...

Orcanopterus is a genus of a eurypterid classified as part of the family Waeringopteridae. The genus contains one species, O. manitoulinensis, from the...

Holmipterus is a problematic genus of eurypterid, an extinct group of aquatic arthropods. The type and only species of Holmipterus, H. suecicus, is known...

we cannot assign this new arthropod to the trilobites or any living chelicerate or crustacean taxon. It represents an intermediate morphology like those...

Parahughmilleria (meaning "near Hughmilleria") is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Parahughmilleria have been...

Vernonopterus is a genus of eurypterid, a group of extinct aquatic arthropods. Fossils of Vernonopterus have been discovered in deposits of the Carboniferous...

Pentlandopterus is a genus of prehistoric eurypterid from the Late Ordovician period. The genus contains one species, P. minor, known from the Pentland...

distinguished from other groups of arthropods, such as insects, myriapods and chelicerates by the possession of biramous (two-parted) limbs, and by the nauplius...

Pseudoniscus is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Pseudoniscus was regarded as part of the clade Planaterga...

Tylopterella is a genus of eurypterid, a group of extinct aquatic arthropods. Only one fossil of the single and type species, T. boylei, has been discovered...

Tarsopterella is a genus of prehistoric eurypterid classified within the family Hardieopteridae. It contains only one species, T. brewsteri from the Lower...

relative length of the leg segments, small male palpus (appendages of chelicerates), and the weakly sclerotized (hardened) epigynum in females. Metepeira...

of Kasibelinurus; Selden, Lamsdell & Qi (2015) reinterpreted it as a chelicerate belonging to the lineage that branched from the main euchelicerate lineage...

helped solidify the close relationship between the eurypterids and other chelicerates by showcasing numerous homologies between the two groups. Samuel Almond...

red-tipped halfbeak, banded archerfish, and spotted green pufferfish Chelicerates: horseshoe crab Crustaceans: fiddler crab, mangrove hermit crab, and...

distinguished from other groups of arthropods, such as insects, myriapods and chelicerates, by the possession of biramous (two-parted) limbs, and by their larval...

(2018). "The gnathobasic spine microstructure of recent and Silurian chelicerates and the Cambrian artiopodan Sidneyia: Functional and evolutionary implications"...

Necrogammarus salweyi is the binomial name applied to an arthropod fossil discovered in Herefordshire, England. The fossil represents a fragmentary section...

Skabelund; Javier Ortega-Hernández (2020). "Revision of the mollisoniid chelicerate(?) Thelxiope, with a new species from the middle Cambrian Wheeler Formation...

other living arthropods, their traditional classification as a member of chelicerates alongside horseshoe crabs and arachnids has regained wide support in...

Caron (2017). "Mandibulate convergence in an armoured Cambrian stem chelicerate". BMC Evolutionary Biology. 17 (1) 261. doi:10.1186/s12862-017-1088-7...

Lamsdell & Qi Devonian (Lochkovian) Xiaxishancun Formation China A chelicerate belonging to the lineage that branched from the main euchelicerate lineage...

Cambrian mollisoniid and the ancestral nervous system organization of chelicerates". Nature Communications. 13 (1) 410. Bibcode:2022NatCo..13..410O. doi:10...