Adocus

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Adocus Temporal range: Early Cretaceous - Oligocene, 125–28 Ma PreꞒ Ꞓ O S D C P T J K Pg N

Shell of Adocus beatus, Peabody Museum of Natural History
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Family:	†Adocidae
Subfamily:	†Adocinae
Genus:	†Adocus Cope, 1868
Type species
†Adocus beatus Leidy, 1865
Species
See text

Adocus is an extinct genus of aquatic turtles belonging to the family Adocidae.

Description

Species of the genus Adocus had flattened and smoothly contoured shells with horny sculptured plates. The shells could reach a length of at least 50 centimetres (1.6 ft) for North American species, some species like A. kohaku had carapace length of 60 centimetres (2.0 ft).^[1] The largest species, A. kirtlandius had carapace reaching 113 centimetres (3.71 ft).^[2] These large freshwater turtles had an omnivorous diet. They lived from the Late Cretaceous to the Paleocene in North America, but in Asia, they were also present during the Oligocene.

Distribution

These turtles have been found in Cretaceous to Paleogene of Canada, United States, Mongolia, China, Japan, Kazakhstan, Tajikistan, and Uzbekistan.^[3]

Species

Adocus agilis
Adocus aksary
Adocus beatus, type species (synonyms: A. punctatus, A. lacer)
Adocus bossi
Adocus bostobensis
Adocus dzhurtasensis
Adocus firmus
Adocus foveatus
Adocus hesperius
Adocus kirtlandius
Adocus kizylkumensis
Adocus kohaku [ja]^[1]
Adocus lineolatus
Adocus onerosus
Adocus orientalis
Adocus pravus
Adocus sengokuensis [ja]^[4]
Adocus syntheticus

References

^ ^a ^b Hirayama*, Ren; Sonoda, Teppei; Uno, Hikaru; Horie, Kenji; Tsutsumi, Yukiyasu; Sasaki, Kazuhisa; Takisawa, Shunsuke Mitsuzuka and Toshio (2022-03-14). "Adocus Kohaku, A New Species of Aquatic Turtle (Testudines: Cryptodira: Adocidae) from the Late Cretaceous of Kuji, Iwate Prefecture, Northeast Japan, with Special References to the Geological Age of the Tamagawa Formation (Kuji Group)". International Journal of Paleobiology & Paleontology. 4 (1).
^ Farina, Bruna M.; Godoy, Pedro L.; Benson, Roger B. J.; Langer, Max C.; Ferreira, Gabriel S. (2023). "Turtle body size evolution is determined by lineage-specific specializations rather than global trends". Ecology and Evolution. 13 (6) e10201. Bibcode:2023EcoEv..1310201F. doi:10.1002/ece3.10201. ISSN 2045-7758. PMC 10293707. PMID 37384241.
^ "福岡県からの白亜紀前期の新種のカメ化石について". 福井県立恐竜博物館 (in Japanese). Retrieved 2023-11-27.
^ Teppei Sonoda; Ren Hirayama; Yoshihiko Okazaki; Hisao Ando (2015). "A New Species of the Genus Adocus (Adocidae, Testudines) from the Lower Cretaceous of Southwest Japan". Paleontological Research. 19 (1): 26–32. doi:10.2517/2014PR026. S2CID 130008038.

The Paleobiology Database Archived 2007-10-01 at the Wayback Machine
Paleocene Mammals
Recently Collected Specimen of Adocus
E.V. Syromyatnikova and I.G. Danilov [1]
Yale Digital Content

Testudines

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Diapsida
Order: Testudines

Genus

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Gigantochelys †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Kinosternoidea

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Testudinoidea

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Trionychia

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

Pleurodira

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based on Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.

See also List of Testudines families

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